The base composition patterns of genes, coding sequences and gene expression levels were analyzed in the available long sequences (contigs) of Arabidopsis. Chromosome 5 was analyzed in detail and all chromosomes for which sequence data are now available show essentially the same large-scale compositional properties. Guanine+cytosine levels of genes and of their coding regions, as well as gene densities and expression levels, all show a marked tendency to be higher in the distal regions of Arabidopsis chromosomes.